Introduction to the Mollusca Phylum

The Mollusca phylum encompasses a vast diversity of invertebrates, including snails, clams, and cephalopods like squids and octopuses. These creatures are characterized by a muscular foot, mantle, and often a radula. Originating over 500 million years ago, molluscs have a rich fossil record and play significant roles in human economies and ecosystems. Conservation efforts are crucial for nearly 2,000 endangered non-marine species.

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Introduction to the Mollusca Phylum

The Mollusca phylum, second only to Arthropoda in terms of invertebrate diversity, includes a vast array of organisms collectively known as molluscs. With around 85,000 extant species and an estimated 60,000 to 100,000 fossil species, molluscs are a significant presence in marine environments, comprising about 23% of marine organisms. They also thrive in freshwater and on land. Molluscs display remarkable diversity in size, form, behavior, and habitat. The phylum is categorized into numerous classes, including Gastropoda, Bivalvia, Cephalopoda, and others, with Gastropoda alone accounting for 80% of living molluscs. Cephalopods, such as squids, octopuses, and cuttlefish, are particularly notable for their complex nervous systems.
Variety of live molluscs on a neutral background: spiral gastropod, blue and purple bivalves, elongated cephalopod and small snails.

Distinctive Features of Molluscs

Molluscs are unified by certain anatomical features: a muscular foot for locomotion, a mantle which secretes the shell and forms a respiratory cavity, and, in most classes, a radula for feeding. Their body structure varies greatly, leading to the concept of a "hypothetical ancestral mollusc" to illustrate common traits. This ancestral form is thought to have had a shell composed of protein and chitin, strengthened with calcium carbonate, and a mantle covering its dorsal side. The ventral side would have featured the muscular foot. Molluscs typically have a reduced coelom, with the primary body cavity being a hemocoel that accommodates an open circulatory system. Their nervous system includes paired nerve cords and often a brain, and they possess sensory organs for environmental detection. Reproductive strategies among molluscs are diverse, with many species laying eggs that develop into larvae or miniature adults.

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1

Mollusca phylum rank in invertebrate diversity

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Second to Arthropoda in species variety among invertebrates.

2

Number of extant and fossil mollusc species

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Approximately 85,000 living species, 60,000 to 100,000 fossil species.

3

Molluscs' habitat diversity

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Inhabit marine, freshwater, and terrestrial environments.

4

The concept of a '______ ancestral mollusc' is used to demonstrate shared characteristics, which is believed to have had a shell made of ______ and ______, reinforced with ______.

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hypothetical protein chitin calcium carbonate

5

Molluscs generally have a limited ______, with their main body cavity being a ______ that houses an ______ circulatory system.

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coelom hemocoel open

6

Evolutionary debate of molluscs' origin

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Molluscs' origins are contested; emerged in Cambrian, details unclear.

7

Molluscs' transition from ancestors

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Evolved from Lophotrochozoans; research ongoing on diversity development.

8

The shells of ______ have historically been used as ______.

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molluscs currency

9

Certain ______ like the blue-ringed octopus and some cone snails are known for their ______.

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molluscs venom

10

Molluscs can be problematic as ______ pests or ______ species.

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agricultural invasive

11

Some freshwater snails act as hosts for a parasite that causes ______, impacting millions worldwide.

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schistosomiasis

12

Origin of 'mollusc' term

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Derived from Latin 'mollusca' meaning 'soft'; first used in 17th century.

13

Obsolete classification 'Molluscoida'

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Former grouping of unrelated animals with molluscs; discarded due to phylogenetic advances.

14

______ display a wide variety of shapes and special features, including well-known ______ and ______.

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Molluscs snails clams

15

These creatures can be found in ______, ______, and ______ environments.

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marine freshwater terrestrial

16

The greatest variety of these species is observed in ______ and ______ regions.

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tropical temperate

17

Many species that do not live in the ocean are at risk of ______, with nearly ______ listed as endangered by the ______.

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extinction 2,000 IUCN Red List

18

To maintain the biological diversity and ecological roles of these organisms, ______ ______ are crucial.

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conservation efforts

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