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Introduction to the Mollusca Phylum

The Mollusca phylum encompasses a vast diversity of invertebrates, including snails, clams, and cephalopods like squids and octopuses. These creatures are characterized by a muscular foot, mantle, and often a radula. Originating over 500 million years ago, molluscs have a rich fossil record and play significant roles in human economies and ecosystems. Conservation efforts are crucial for nearly 2,000 endangered non-marine species.

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1

Mollusca phylum rank in invertebrate diversity

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Second to Arthropoda in species variety among invertebrates.

2

Number of extant and fossil mollusc species

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Approximately 85,000 living species, 60,000 to 100,000 fossil species.

3

Molluscs' habitat diversity

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Inhabit marine, freshwater, and terrestrial environments.

4

The concept of a '______ ancestral mollusc' is used to demonstrate shared characteristics, which is believed to have had a shell made of ______ and ______, reinforced with ______.

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hypothetical protein chitin calcium carbonate

5

Molluscs generally have a limited ______, with their main body cavity being a ______ that houses an ______ circulatory system.

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coelom hemocoel open

6

Evolutionary debate of molluscs' origin

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Molluscs' origins are contested; emerged in Cambrian, details unclear.

7

Molluscs' transition from ancestors

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Evolved from Lophotrochozoans; research ongoing on diversity development.

8

The shells of ______ have historically been used as ______.

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molluscs currency

9

Certain ______ like the blue-ringed octopus and some cone snails are known for their ______.

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molluscs venom

10

Molluscs can be problematic as ______ pests or ______ species.

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agricultural invasive

11

Some freshwater snails act as hosts for a parasite that causes ______, impacting millions worldwide.

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schistosomiasis

12

Origin of 'mollusc' term

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Derived from Latin 'mollusca' meaning 'soft'; first used in 17th century.

13

Obsolete classification 'Molluscoida'

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Former grouping of unrelated animals with molluscs; discarded due to phylogenetic advances.

14

______ display a wide variety of shapes and special features, including well-known ______ and ______.

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Molluscs snails clams

15

These creatures can be found in ______, ______, and ______ environments.

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marine freshwater terrestrial

16

The greatest variety of these species is observed in ______ and ______ regions.

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tropical temperate

17

Many species that do not live in the ocean are at risk of ______, with nearly ______ listed as endangered by the ______.

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extinction 2,000 IUCN Red List

18

To maintain the biological diversity and ecological roles of these organisms, ______ ______ are crucial.

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conservation efforts

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Introduction to the Mollusca Phylum

The Mollusca phylum, second only to Arthropoda in terms of invertebrate diversity, includes a vast array of organisms collectively known as molluscs. With around 85,000 extant species and an estimated 60,000 to 100,000 fossil species, molluscs are a significant presence in marine environments, comprising about 23% of marine organisms. They also thrive in freshwater and on land. Molluscs display remarkable diversity in size, form, behavior, and habitat. The phylum is categorized into numerous classes, including Gastropoda, Bivalvia, Cephalopoda, and others, with Gastropoda alone accounting for 80% of living molluscs. Cephalopods, such as squids, octopuses, and cuttlefish, are particularly notable for their complex nervous systems.
Variety of live molluscs on a neutral background: spiral gastropod, blue and purple bivalves, elongated cephalopod and small snails.

Distinctive Features of Molluscs

Molluscs are unified by certain anatomical features: a muscular foot for locomotion, a mantle which secretes the shell and forms a respiratory cavity, and, in most classes, a radula for feeding. Their body structure varies greatly, leading to the concept of a "hypothetical ancestral mollusc" to illustrate common traits. This ancestral form is thought to have had a shell composed of protein and chitin, strengthened with calcium carbonate, and a mantle covering its dorsal side. The ventral side would have featured the muscular foot. Molluscs typically have a reduced coelom, with the primary body cavity being a hemocoel that accommodates an open circulatory system. Their nervous system includes paired nerve cords and often a brain, and they possess sensory organs for environmental detection. Reproductive strategies among molluscs are diverse, with many species laying eggs that develop into larvae or miniature adults.

Molluscan Evolution and the Fossil Record

The evolutionary origins of molluscs are debated, but it is generally accepted that they emerged during the Cambrian period, over 500 million years ago. The transition from their Lophotrochozoan ancestors to the present-day diversity is a topic of ongoing research. The fossil record, including well-preserved specimens like ammonites, provides valuable insights into the phylum's ancient diversity and the environmental conditions of past eras.

Molluscs and Human Interaction

Molluscs have been integral to human diets and economies, providing food, pearls, mother of pearl, and other luxury items. Their shells have even functioned as currency. However, some species pose risks; for instance, the blue-ringed octopus and certain cone snails are venomous. Molluscs can also be agricultural pests or invasive species, disrupting local ecosystems. Additionally, some freshwater snails are intermediate hosts for the parasite causing schistosomiasis, a disease affecting millions globally.

The Study of Molluscs: Malacology

The term "mollusc" originates from the Latin "mollusca," meaning "soft," which was first applied to these animals in the 17th century. Malacology is the branch of zoology that focuses on the study of molluscs. The outdated term "Molluscoida," which previously grouped unrelated animals such as brachiopods and tunicates with molluscs, has been discarded due to advances in phylogenetic understanding.

Molluscan Diversity and Conservation

Molluscs exhibit a vast range of forms and adaptations, from the familiar snails and clams to more obscure groups. They inhabit diverse ecosystems, including marine, freshwater, and terrestrial habitats. The highest diversity is found in tropical and temperate zones. Despite their prevalence, many non-marine molluscs face extinction threats, with the IUCN Red List citing nearly 2,000 endangered species. Conservation efforts are essential to preserve these species and their ecological contributions.